Leptotarsus

Leptotarsus
Leptotarsus viridis
Nomenklatur
Leptotarsus Guérin-Ménéville, 1831
Populærnavn
storstankelbein, stankelbein
Klassifikasjon
Rike		Dyreriket
Rekke		Leddyr
Klasse		Insekter
Orden		Tovinger
Underorden		Mygg
Familie		Storstankelbein
Økologi
Antall arter:		ca. 300
Utbredelse:		varme områder, ikke i Europa
Inndelt i

Leptotarsus er en artsrik slekt av stankelbein, er storvoksne mygg som hører til familien storstankelbein (Tipulidae).

Utseende

Store, langbeinte, vanligvis lyst fargede (gulaktige, rødlige, grønnlige eller lysbrune) stankelbein. Antennene er trådformede. Mange har fargetegninger på vingene.

Levevis

Larvene lever i jorden eller mellom råtnende plantemateriale.

En gammel slekt

Fossiler av denne slekten er kjent tilbake til nedre kritt ^[1], slekten er dermed en av de eldste kjente i Tipulidae.

Utbredelse

De fleste av artene lever i tropiske områder på den sørlige halvkulen (rundt halvparten i Australia og en tredjedel i Sør-Amerika), men slekten forekommer så langt nord som Nord-Amerika. Den finnes ikke i Europa. Slekten erstatter på mange måter slekten Tipula på den sørlige halvkulen.

Systematisk inndeling

Listen følger ^[2].

Treliste

Orden tovinger (Diptera)
- Underorden mygg (Nematocera)
  - Overfamilien stankelbein (Tipuloidea)
    - Familien storstankelbein (Tipulidae)
      - Slekten Leptotarsus Guérin-Ménéville, 1831
        Underslekten Aeshnasoma Johnson, 1909
        Leptotarsus rivertonensis (Johnson, 1909)
        
        Underslekten Araucomyia Alexander, 1929
        Leptotarsus brevihirsutus (Alexander, 1934)
        
        Leptotarsus chilensis (Philippi, 1866)
        
        Leptotarsus paulseni (Philippi, 1866)
        
        Leptotarsus penitus (Alexander, 1944)
        
        Underslekten Aurotipula Alexander, 1924
        Leptotarsus apertus (Edwards, 1923)
        
        Leptotarsus atroflavus (Alexander, 1922)
        
        Leptotarsus auroatrus (Edwards, 1923)
        
        Leptotarsus bivittatus (Edwards, 1923)
        
        Leptotarsus brevitarsis (Edwards, 1923)
        
        Leptotarsus clarus (Kirby, 1884)
        
        Leptotarsus dux (Kirby, 1884)
        
        Leptotarsus ferruginosus (Edwards, 1923)
        
        Leptotarsus flavoscapus (Alexander, 1922)
        
        Leptotarsus neali Oosterbroek, 1989
        
        Leptotarsus occlusus (Edwards, 1924)
        
        Leptotarsus orion (Hudson, 1895)
        
        Leptotarsus ruapehuensis (Alexander, 1923)
        
        Leptotarsus subtener (Alexander, 1922)
        
        Leptotarsus tener (Hutton, 1900)
        
        Underslekten Brevicera Miller, 1945
        Leptotarsus aenigmaticus (Alexander, 1926)
        
        Leptotarsus heterogamus (Hudson, 1913)
        
        Leptotarsus waitakerensis (Alexander, 1952)
        
        Underslekten Ceoneura Alexander, 1924
        Leptotarsus idioneura (Alexander, 1924)
        
        Underslekten Chlorotipula Alexander, 1924
        Leptotarsus albistigma (Edwards, 1923)
        
        Leptotarsus angustior (Alexander, 1923)
        
        Leptotarsus elongatus (Edwards, 1923)
        
        Leptotarsus holochlorus (Nowicki, 1875)
        
        Leptotarsus virescens (Edwards, 1923)
        
        Leptotarsus viridis (Walker, 1856)
        
        Underslekten Habromastix Skuse, 1890
        Leptotarsus bulburinensis Dobrotworsky, 1974
        
        Leptotarsus cinerascens (Skuse, 1890)
        
        Leptotarsus cunninghamensis Dobrotworsky, 1974
        
        Leptotarsus heroni (Alexander, 1922)
        
        Leptotarsus hilli (Alexander, 1922)
        
        Leptotarsus luciae (Alexander, 1948)
        
        Leptotarsus novellus (Alexander, 1928)
        
        Leptotarsus ornatipes (Skuse, 1890)
        
        Leptotarsus parallelus (Alexander, 1920)
        
        Leptotarsus pergrandis (Alexander, 1922)
        
        Leptotarsus platyxanthus (Alexander, 1924)
        
        Leptotarsus remotus (Walker, 1848)
        
        Leptotarsus robinsoni (Alexander, 1924)
        
        Leptotarsus similior (Alexander, 1923)
        
        Leptotarsus sublateralis (Alexander, 1922)
        
        Leptotarsus terrareginae (Alexander, 1920)
        
        Underslekten Kocia Kocak & Kemal, 2009
        Leptotarsus gesneri (Enderlein, 1912)
        
        Underslekten Leptotarsus (i snever forstand)
        Leptotarsus annulipes (Philippi, 1866)
        
        Leptotarsus ardrossanensis Dobrotworsky, 1972
        
        Leptotarsus bancrofti (Alexander, 1929)
        
        Leptotarsus caesareus Alexander, 1929
        
        Leptotarsus cinereipleura Alexander, 1929
        
        Leptotarsus clavatus (Maquart, 1850)
        
        Leptotarsus coolgardiensis Dobrotworsky, 1972
        
        Leptotarsus ducalis (Westwood, 1876)
        
        Leptotarsus esperanceiensis Dobrotworsky, 1972
        
        Leptotarsus flavolateralis Alexander, 1944
        
        Leptotarsus fletcherensis Dobrotworsky, 1972
        
        Leptotarsus fraucai Dobrotworsky, 1972
        
        Leptotarsus gloria (Alexander, 1929)
        
        Leptotarsus imperatorius (Westwood, 1876)
        
        Leptotarsus kalamundaensis Dobrotworsky, 1972
        
        Leptotarsus macquarti Guérin-Ménéville, 1831
        
        Leptotarsus nigrithorax (Macquart, 1850)
        
        Leptotarsus regificus (Alexander, 1922)
        
        Leptotarsus rufosternatus Alexander, 1944
        
        Leptotarsus scutellaris Skuse, 1890
        
        Leptotarsus subapterus Alexander, 1978
        
        Leptotarsus tricinctus (Walker, 1848)
        
        Leptotarsus trivittatus Skuse, 1890
        
        Underslekten Leptotipula Alexander, 1917
        Leptotarsus edwardsianus (Alexander, 1920)
        
        Leptotarsus limnophiloides (Alexander, 1917)
        
        Underslekten Limoniodes Alexander, 1938
        Leptotarsus sulphurellus (Alexander, 1938)
        
        Underslekten Longurio Loew, 1869
        Leptotarsus africanus (Alexander, 1921)
        
        Leptotarsus albicubitalis (Alexander, 1964)
        
        Leptotarsus anoplostylus (Alexander, 1964)
        
        Leptotarsus aspropodus (Alexander, 1957)
        
        Leptotarsus atrirostris (Alexander, 1945)
        
        Leptotarsus basuticanus (Alexander, 1956)
        
        Leptotarsus belloides (Alexander, 1921)
        
        Leptotarsus bertii Alexander, 1979
        
        Leptotarsus bonaespaei (Bergroth, 1888)
        
        Leptotarsus borgmeieranus Alexander, 1969
        
        Leptotarsus brasiliae (Alexander, 1935)
        
        Leptotarsus browni (Alexander, 1940)
        
        Leptotarsus bullocki (Alexander, 1931)
        
        Leptotarsus byersi Young & Gelhaus, 1992
        
        Leptotarsus caffer (Alexander, 1917)
        
        Leptotarsus caparaonus (Alexander, 1944)
        
        Leptotarsus capicola (Alexander, 1921)
        
        Leptotarsus carreranus (Alexander, 1945)
        
        Leptotarsus chaoianus (Alexander, 1949)
        
        Leptotarsus chionoides (Alexander, 1917)
        
        Leptotarsus chrysostigma (Alexander, 1944)
        
        Leptotarsus cinereilinea (Alexander, 1921)
        
        Leptotarsus cnephosus (Alexander, 1960)
        
        Leptotarsus congestus (Alexander, 1949)
        
        Leptotarsus coronatus (Alexander, 1917)
        
        Leptotarsus dolichoros (Wood, 1952)
        
        Leptotarsus drakensbergensis (Alexander, 1956)
        
        Leptotarsus eshowensis (Alexander, 1960)
        
        Leptotarsus espinozai (Alexander, 1939)
        
        Leptotarsus eucryptus (Alexander, 1953)
        
        Leptotarsus exemptus (Alexander, 1936)
        
        Leptotarsus flagellatus (Wood, 1952)
        
        Leptotarsus fulvus (Edwards, 1916)
        
        Leptotarsus ganocephalus (Alexander, 1960)
        
        Leptotarsus goyazanus (Alexander, 1940)
        
        Leptotarsus giumaraesi (Alexander, 1942)
        
        Leptotarsus gurneyi Alexander, 1975
        
        Leptotarsus gymnocerus (Alexander, 1938)
        
        Leptotarsus hainanensis (Alexander, 1936)
        
        Leptotarsus helotus Alexander, 1969
        
        Leptotarsus hirsutistylus (Alexander, 1949)
        
        Leptotarsus huanucensis (Alexander, 1954)
        
        Leptotarsus inaequipes (Alexander, 1956)
        
        Leptotarsus insidiosus (Alexander, 1949)
        
        Leptotarsus ixion (Alexander, 1949)
        
        Leptotarsus jonesi (Alexander, 1920)
        
        Leptotarsus lemniscatus (Alexander, 1929)
        
        Leptotarsus lustralis (Alexander, 1936)
        
        Leptotarsus luteiniger (Alexander, 1963)
        
        Leptotarsus luteistigma (Alexander, 1940)
        
        Leptotarsus macarius (Alexander, 1963)
        
        Leptotarsus melanopterus (Alexander, 1930)
        
        Leptotarsus micropteryx (Alexander, 1921)
        
        Leptotarsus millotianus (Alexander, 1958)
        
        Leptotarsus minimus (Alexander, 1914)
        
        Leptotarsus minusculoides (Wood, 1952)
        
        Leptotarsus minusculus (Alexander, 1917)
        
        Leptotarsus mitiformis (Alexander, 1960)
        
        Leptotarsus mossambicensis (Alexander, 1920)
        
        Leptotarsus mosselensis (Alexander, 1945)
        
        Leptotarsus nahuelbutae (Alexander, 1945)
        
        Leptotarsus neorinus (Alexander, 1953)
        
        Leptotarsus niphopodus (Alexander, 1956)
        
        Leptotarsus nocivus (Alexander, 1953)
        
        Leptotarsus paraguayanus (Riedel, 1921)
        
        Leptotarsus paraguayensis (Alexander, 1935)
        
        Leptotarsus perglabratus (Alexander, 1954)
        
        Leptotarsus phaedrus (Alexander, 1949)
        
        Leptotarsus piger (Alexander, 1945)
        
        Leptotarsus pruinosus (Johnson, 1913)
        
        Leptotarsus pulverosus (Matsumura, 1916)
        
        Leptotarsus pygmaeus (Alexander, 1914)
        
        Leptotarsus quadriniger (Alexander, 1949)
        
        Leptotarsus rabelloi (Alexander, 1954)
        
        Leptotarsus rhodesiae (Alexander, 1937)
        
        Leptotarsus riedelianus (Alexander, 1920)
        
        Leptotarsus rubriceps (Edwards, 1916)
        
        Leptotarsus rubroniger (Alexander, 1921)
        
        Leptotarsus serotinellus (Alexander, 1928)
        
        Leptotarsus sessoris (Alexander, 1954)
        
        Leptotarsus silvester (Wood, 1952)
        
        Leptotarsus spinosus (Wood, 1952)
        
        Leptotarsus stenodiastema (Alexander, 1961)
        
        Leptotarsus stenostylus (Alexander, 1953)
        
        Leptotarsus stuckenbergi (Alexander, 1956)
        
        Leptotarsus styx (Alexander, 1954)
        
        Leptotarsus syndactylus (Alexander, 1956)
        
        Leptotarsus testaceus (Loew, 1869) - forekommer i Nord-Amerika
        
        Leptotarsus tijucanus (Alexander, 1943)
        
        Leptotarsus tinctorius (Alexander, 1960)
        
        Leptotarsus travassosanus (Alexander, 1942)
        
        Leptotarsus variceps (Alexander, 1935)
        
        Leptotarsus versfeldi (Wood, 1952)
        
        Leptotarsus vulsurus (Alexander, 1955)
        
        Leptotarsus yanoi (Alexander, 1953)
        
        Leptotarsus zeylanicus (Alexander, 1958)
        
        Leptotarsus zikanellus (Alexander, 1954)
        
        Underslekten Macromastix Osten Sacken, 1887
        Leptotarsus albicollis (Alexander, 1924)
        
        Leptotarsus albipedis Hynes, 1993
        
        Leptotarsus albiplagius (Alexander, 1923)
        
        Leptotarsus alexanderi (Edwards, 1923)
        
        Leptotarsus alfie Theischinger, 1996
        
        Leptotarsus amissionis (Alexander, 1952)
        
        Leptotarsus angusticosta (Alexander, 1923)
        
        Leptotarsus arenarius (Alexander, 1950)
        
        Leptotarsus atridorsum (Alexander, 1922)
        
        Leptotarsus aurantioceps (Alexander, 1924)
        
        Leptotarsus barringtoniensis Dobrotworsky, 1974
        
        Leptotarsus binnaburrae (Alexander, 1951)
        
        Leptotarsus binotatus (Hutton, 1900)
        
        Leptotarsus brachycerus Alexander, 1978
        
        Leptotarsus brevipetiolatus (Alexander, 1922)
        
        Leptotarsus brisbaneiensis Dobrotworsky, 1974
        
        Leptotarsus caledonianus (Alexander, 1934)
        
        Leptotarsus campbelli (Alexander, 1923)
        
        Leptotarsus cinereus (Edwards, 1923)
        
        Leptotarsus clarkianus (Alexander, 1930)
        
        Leptotarsus clitellarius (Alexander, 1930)
        
        Leptotarsus cockerellae (Alexander, 1929)
        
        Leptotarsus collessi Dobrotworsky, 1974
        
        Leptotarsus constrictus (Skuse, 1890)
        
        Leptotarsus cubitalis (Edwards, 1923)
        
        Leptotarsus decoratus (Edwards, 1923)
        
        Leptotarsus dichroithorax (Alexander, 1920)
        
        Leptotarsus dimidiatus (Westwood, 1835)
        
        Leptotarsus dispar (Walker, 1835)
        
        Leptotarsus dorrigensis (Alexander, 1924)
        
        Leptotarsus errans (Edwards, 1927)
        
        Leptotarsus fergusoni (Alexander, 1924)
        
        Leptotarsus flavidipennis (Alexander, 1923)
        
        Leptotarsus fucatus (Hutton, 1900)
        
        Leptotarsus fulvithorax Alexander, 1978
        
        Leptotarsus fumibasis (Edwards, 1923)
        
        Leptotarsus fuscolateratus (Alexander, 1922)
        
        Leptotarsus gargettensis Dobrotworsky, 1974
        
        Leptotarsus glabristylus Hynes, 1993
        
        Leptotarsus glaucocapillus (Alexander, 1952)
        
        Leptotarsus greyanus (Alexander, 1922)
        
        Leptotarsus hackeri (Alexander, 1920)
        
        Leptotarsus halteratus (Alexander, 1923)
        
        Leptotarsus harrisi (Alexander, 1922)
        
        Leptotarsus helmsi (Skuse, 1890)
        
        Leptotarsus hudsonianus (Alexander, 1922)
        
        Leptotarsus humilis (Skuse, 1890)
        
        Leptotarsus huttoni (Edwards, 1923)
        
        Leptotarsus igniceps (Alexander, 1928)
        
        Leptotarsus incertus (Edwards, 1923)
        
        Leptotarsus intermedius (Alexander, 1922)
        
        Leptotarsus limbipennis (Macquart, 1838)
        
        Leptotarsus longioricornis (Alexander, 1923)
        
        Leptotarsus lunatus (Hutton, 1900)
        
        Leptotarsus luteicosta (Alexander, 1924)
        
        Leptotarsus luteisubcostatus (Alexander, 1934)
        
        Leptotarsus macquartianus (Alexander, 1924)
        
        Leptotarsus mastersi (Skuse, 1890)
        
        Leptotarsus mathewsi (Alexander, 1930)
        
        Leptotarsus mesocerus (Alexander, 1922)
        
        Leptotarsus minor (Edwards, 1923)
        
        Leptotarsus minutissimus (Alexander, 1923)
        
        Leptotarsus mixtus Hynes, 1993
        
        Leptotarsus monstratus (Alexander, 1924)
        
        Leptotarsus montanus (Hutton, 1900)
        
        Leptotarsus mouicola (Alexander, 1951)
        
        Leptotarsus mutabilis (Alexander, 1928)
        
        Leptotarsus nigropolitus (Alexander, 1928)
        
        Leptotarsus noelianus Alexander, 1978
        
        Leptotarsus novocaledonicus (Alexander, 1929)
        
        Leptotarsus obliquus (Edwards, 1923)
        
        Leptotarsus obliteratus (Alexander, 1923)
        
        Leptotarsus obscuripennis (Kirby, 1884)
        
        Leptotarsus obscurirostris (Skuse, 1890)
        
        Leptotarsus ohakunensis (Alexander, 1923)
        
        Leptotarsus opifex (Alexander, 1929)
        
        Leptotarsus ornatus (Jaennicke, 1867)
        
        Leptotarsus pacificus (Erichson, 1842)
        
        Leptotarsus pallidistigma (Alexander, 1922)
        
        Leptotarsus pallidus (Hutton, 1900)
        
        Leptotarsus pedestris (Alexander, 1939)
        
        Leptotarsus productifrons (Alexander, 1945)
        
        Leptotarsus pseudoaurantioceps Dobrotworsky, 1974
        
        Leptotarsus pseudotortilis Dobrotworsky, 1974
        
        Leptotarsus repletus (Alexander, 1948)
        
        Leptotarsus risbeci (Alexander, 1934)
        
        Leptotarsus rufibasis (Alexander, 1922)
        
        Leptotarsus rufiventris (Edwards, 1923)
        
        Leptotarsus sessilis (Alexander, 1924)
        
        Leptotarsus setivena (Alexander, 1929)
        
        Leptotarsus simillimus (Alexander, 1924)
        
        Leptotarsus sinclairi (Edwards, 1923)
        
        Leptotarsus spinastylus Hynes, 1993
        
        Leptotarsus submancus (Alexander, 1923)
        
        Leptotarsus submontanus (Edwards, 1923)
        
        Leptotarsus subnitidus (Alexander, 1924)
        
        Leptotarsus subobsoletus (Alexander, 1926)
        
        Leptotarsus subvittatus (Alexander, 1939)
        
        Leptotarsus tamborineiensis Dobrotworsky, 1974
        
        Leptotarsus tapleyi (Alexander, 1923)
        
        Leptotarsus tenuifrons (Alexander, 1926)
        
        Leptotarsus tortilis (Alexander, 1920)
        
        Leptotarsus variegatus (Edwards, 1923)
        
        Leptotarsus verreauxi (Alexander, 1929)
        
        Leptotarsus vittatus (Edwards, 1923)
        
        Leptotarsus vulpinus (Hutton, 1881)
        
        Leptotarsus zeylandiae (Alexander, 1920)
        
        Underslekten Maoritipula Alexander, 1924
        Leptotarsus hudsonellus (Alexander, 1964)
        
        Leptotarsus hudsoni (Alexander, 1924)
        
        Leptotarsus maori (Alexander, 1920)
        
        Leptotarsus nebulosus (Hudson, 1920)
        
        Underslekten Pehlkea Enderlein, 1912
        Leptotarsus columbianus (Enderlein, 1912)
        
        Leptotarsus pallitarsis (Alexander, 1937)
        
        Leptotarsus regina (Alexander, 1914)
        
        Leptotarsus regulus (Alexander, 1943)
        
        Underslekten Phymatopsis Skuse, 1890
        Leptotarsus albidipes (Alexander, 1929)
        
        Leptotarsus brevipalpis (Alexander, 1920)
        
        Leptotarsus brevirostratus (Alexander, 1922)
        
        Leptotarsus flavopygialis (Alexander, 1920)
        
        Leptotarsus intricatus (Alexander, 1928)
        
        Leptotarsus nigrirostris (Skuse, 1890)
        
        Leptotarsus nigrolimbatus (Alexander, 1929)
        
        Leptotarsus nigrosubcostatus (Alexander, 1924)
        
        Leptotarsus periplocus (Alexander, 1928)
        
        Leptotarsus tenuirostris (Alexander, 1924)
        
        Leptotarsus tonnoiranus (Alexander, 1928)
        
        Leptotarsus walpoleiensis Dobrotworsky, 1974
        
        Underslekten Pseudoleptotarsus Alexander, 1920
        Leptotarsus liponeurus Alexander, 1920
        
        Leptotarsus nigrinus Dobrotworsky, 1974
        
        Underslekten Tanypremna Osten Sacken, 1886
        Leptotarsus albobasalis Alexander, 1969
        
        Leptotarsus aurantiothorax (Alexander, 1954)
        
        Leptotarsus bezzianus (Alexander, 1921)
        
        Leptotarsus borgmeieri (Alexander, 1942)
        
        Leptotarsus brasiliensis Oosterbroek, 2009
        
        Leptotarsus calliope (Alexander, 1945)
        
        Leptotarsus carbonipes (Alexander, 1938)
        
        Leptotarsus cerritus (Alexander, 1945)
        
        Leptotarsus clotho (Alexander, 1944)
        
        Leptotarsus elegantior (Alexander, 1945)
        
        Leptotarsus fieldianus (Alexander, 1956)
        
        Leptotarsus fuscitarsis (Alexander, 1919)
        
        Leptotarsus gouadeloupensis Young, 2001
        
        Leptotarsus hodgei (Alexander, 1939)
        
        Leptotarsus horridus (Alexander, 1941)
        
        Leptotarsus incompletus (Alexander, 1944)
        
        Leptotarsus invaripes (Alexander, 1936)
        
        Leptotarsus kadeni (Alexander, 1941)
        
        Leptotarsus limai (Alexander, 1945)
        
        Leptotarsus longipes (Fabricius, 1805)
        
        Leptotarsus longissimus (Enderlein, 1912)
        
        Leptotarsus manicatus (Osten Sacken, 1886)
        
        Leptotarsus mirandus (Alexander, 1944)
        
        Leptotarsus opilio (Osten Sacken, 1886)
        
        Leptotarsus ornatipes (Alexander, 1941)
        
        Leptotarsus perornatus (Alexander, 1929)
        
        Leptotarsus phylax (Alexander, 1954)
        
        Leptotarsus picturellus (Alexander, 1941)
        
        Leptotarsus porterianus (Alexander, 1937)
        
        Leptotarsus proavitus (Alexander, 1940)
        
        Leptotarsus salome (Alexander, 1945)
        
        Leptotarsus saltatrix (Alexander, 1944)
        
        Leptotarsus sanctaecatharinae (Alexander, 1945)
        
        Leptotarsus uniguttatus (Alexander, 1954)
        
        Underslekten Tanypremnella Alexander, 1938
        Leptotarsus abitaguensis Oosterbroek, 2009
        
        Leptotarsus antennifer (Alexander, 1942)
        
        Leptotarsus crystallinus (Alexander, 1938)
        
        Leptotarsus elongatus Alexander, 1978
        
        Leptotarsus gentilis (Alexander, 1943)
        
        Leptotarsus maldonadoi (Alexander, 1953)
        
        Leptotarsus mediocornis (Alexander, 1944)
        
        Leptotarsus megacerus (Alexander, 1943)
        
        Leptotarsus microcerus (Alexander, 1943)
        
        Leptotarsus perdistinctus (Alexander, 1942)
        
        Leptotarsus segnipes (Alexander, 1953)
        
        Leptotarsus transfasciatus (Alexander, 1938)
        
        Underslekten Tanypremnodes Alexander, 1924
        Leptotarsus leucoplaca (Alexander, 1924)
        
        Leptotarsus subapicalis (Alexander, 1941)
        
        Underslekten Xenotipula Alexander, 1921
        Leptotarsus cisatlanticus (Alexander, 1937)
        
        Leptotarsus munroi (Alexander, 1921)

Referanser

^ G.C. Ribeiro, D. Santos og R.C. Ramos Nicolau (2015) A new species of Leptotarsus (Diptera: Tipulidae) from the Lower Cretaceous Crato Formation of Brazil. Cretaceous Research 56: 244-249. [1]
^ Oosterbroek, Pjotr (2017): Catalogue of the Craneflies of the World